Botany: 35 families, 13100 species.
Botany: ovary superior, often of 3 fused carpels, dry stigmas; leaves often simple, entire, leathery, fleshy, trilacunar nodes; seed fibrous exotegmen; flowers inconspicuous, reduced, unisexual; ovary of 2 carpels, unilocular, single apical ovule; pollen porate, wind pollinated, stamens 5 or less, single whorl; fruit drupaceous; cystolids of calcium carbonate.
DD: Ericaceae; Fabaceae.
In the Apg3 classification Malpighiales is one of the Orders in Fabidae. Malpighiales is part of the COM clade: Celastrales, Oxalidales and Malpighiales. The order of the Malpighiales is somewhat strange in the sense that it is the one with the least qualities common to all the families. It is based mostly on DNA analysis. A common feature is the trilacunar nodes in the leaf attachment.
Some suborders have formerly been treated as orders such as Violales and Euphorbiales.
In the Cronquist classification Violales included many families, that in the Apg3 classification are transferred to other orders: Cucurbitaceae and Datiscaceae to Cucurbitales, Cistaceae to Malvales, Tamaricaceaea to Caryophyllales and Caricaceae to Brassicales. In the Apg3 classification the “Violales” clade is a Suborder in the Order of the Malpighiales. It includes Violaceae, Passifloraceae, Salicaeae, Turneraceae and Achariaceae.
In the Plant theory the Malpighiales is split into 2 Orders. Violales is split off from Malpighiales. Violales is placed in Phase 3.
1. Rhizophoraceae, Erythroxylaceae, Ctenolophonaceae.
2. Passifloraceae, Turneraceae, Malesherbiaceae.
4. Euphorbiaceae, Peraceae, Picrodendraceea, Phyllantaceae.